Just noticed this.

[Valence Axiom]

Is this an known issue?  Do the allowed enhancements change as you level?  I don't recall seeing this before.

 

I was doing a little enhancement upgrading on a lvl 37 beast/rad mastermind and noticed that the lioness slots have a defense and health enhancement in them.  When I went to throw higher level ones in, it didn't work.  When I look at the allowed enhancement window it shows they're not allowed.  I know they were when I put them in.  It also shows the ones that are there aren't doing anything.  Screenshot shows what I am trying to describe.

[Arbegla]

They recently moved the Defense and Heal enhancements to the equip/upgrade powers instead of the pets themselves. You probably had it slotted before the patch.

[Valence Axiom]

This character has been idle and neglected for a while.  So that's probably it.  Thanks for replying.

 

[Uun]

See patch notes below, particularly the highlighted bits. While they moved slotting of Healing and Resistance enhancements from the summons to Tame Beasts, they eliminated slotting Defense enhancements entirely because they weren't doing anything.

• Fortify Pack
  • This power no longer locks out Pack Mentality stack gain
  • Now has a highlight ring when Pack Mentality stacks reach 10
• Call Ravens
  • Power can now only grant one stack of Pack Mentality
• Personal attacks Call Swarm, Call Hawk, Call Ravens
  • Now grant a 30 second +3% accuracy buff to summoned beasts and the caster when used.
  • Now grant a 30 second +32.128 Max HP buff to summoned beasts and the caster when used.
• Summon Dire Wolf
  • No longer accepts Resistance enhancements or sets
  • No longer accepts Healing enhancements or sets
• Summon Lions
  • No longer accepts Defense Enhancements or sets (no Lioness power ever accepted Defense enhancements).
  • No longer accepts Healing enhancements or sets
• Summon Wolves
  • No longer accepts Healing enhancements
  • No longer accepts ToHit enhancements
• Train Beasts
  • Now Accepts ToHit enhancements and sets
  • No longer has a recharge time
  • No longer grants Lick Wounds to Lioness
  • No longer grants Growl to Alpha Wolf
  • Now grants Ferocious Growl to Lioness
  • Now grants Howl to Howler Wolf
  • Now grants Howl to Alpha Wolf
  • ToHit enhancements in this power should improve ToHit buffs in Howl of Alpha and Howler Wolves
• Tame Beasts
  • Now Accepts Healing and Resistance enhancements and sets
  • No longer has a recharge time
  • No longer grants Howl to Howler Wolf
  • No longer grants Howl to Alpha Wolf
  • No longer grants Ferocious Growl to Lioness
  • Now grants Lick Wounds to Lioness
  • No longer grants Growl to Alpha Wolf
  • Dire Wolf Will of the Wild now can be enhanced for resistance
  • Healing enhancements in this power should improve Healing of Alpha Wolf and Lioness' Lick Wounds
  • Healing enhancements in this power should improve Healing of Dire Wolf's Will of the Wild
  • Resistance enhancements in this power should improve Resistances of Dire Wolf's Will of the Wild